The 51-year-old has overseen success at the Chester-le-Street club, with two Championship victories and the award of an Ashes Test for the venue in 2013.
"I have enjoyed a fantastic 20 years," Harker said.
"The club has changed beyond recognition since I joined the team and it's a real honour to have played a part in the fantastic achievements."
Harker will remain involved with the club as a non-executive director.
Meanwhile, chairman Clive Leach has said the club will take time to consider a replacement following Harker's departure.
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